The rest of Lonchodraco probably looks like this large unnamed ornithocheird

Only the deep toothy jaw tips,
of the pterosaur Lonchodraco giganteus (Hooley 1914; Rodrigues & Kellner 2013; NHMUK PV 39412; originally Pterodactylus giganteus Bowerbank 1846; Fig. 1) are known. Ever wonder what the rest of this pterosaur looked like?

the 174-year wait is over.

Figure 1. Lonchodraco jaw tips. Colors added here.

Figure 1. Lonchodraco jaw tips. Colors added here. For the rest of this genus, see figure 2. The nasal (pink) is laminated between the premaxilla (yellow) and maxilla (green). The jugal (blue) also makes an appearance.

What little is known of Lonchodectes turns out to look like
the (so far) unnamed large ornithocheirid, SMNK PAL 1136 (Fig. 2) one of the largest of all flying pterosaurs. The very few parts they have in common are virtually identical, except for size (note the scale bars provided).

Figure 2. The unnamed giant ornithocheirid, SMNK PAL 1136 has a rostrum quite similar to that of Lonchodectes.

Figure 2. The unnamed giant ornithocheirid, SMNK PAL 1136 has a rostrum quite similar to that of Lonchodectes. With such giant wings, soaring over wave tops would have been ideal, dipping occasionally to feed without getting wet.

As one of the largest flying pterosaurs,

SMNK PAL 1136 (Figs. 2, 3) presents no vestigial terminal wing phalanges. No hyper-elongated neck cervicals are present. This pterosaur was built to soar like a big pelican.

Sorry, giant azhdarchids lovers 
(Fig. 3). Those were not volant, as we learned earlier here. They grew to be so big AFTER they became flightless, like flightless birds do. Giant azhdarchids DO have vestigial wing phalanges and a hyper-elongated neck.

Figure 1. Click to enlarge. The largest flying and non-flying birds and pterosaurs to scale.

Figure 3. Click to enlarge. The largest flying and non-flying birds and pterosaurs to scale.

Earlier workers 
did not match Lonchodraco to the SMNK PAL 1136 specimen. Earlier workers did not name the SMNK specimen. Perhaps someone is working on that specimen at present and other workers are giving him/her the honor/duty of naming it.

Wonder if
the Lonchodraco name will stick to the SMNK specimen?

Recently, Martill et al. 2020 took a close look
at the foramina in the jaw tips of Lonchodraco and thought they indicated enhanced sensitivity of the rostrum tip, which implied tactile feeding. With such giant wings, soaring over wave tops would have been likely, dipping occasionally to feed without getting the wings wet.

Odd that the top workers at the top universities
have decided to spend their time examining tiny pits on a broken 174-year-old pterosaur snout while ignoring the origin of pterosaurs… while ignoring many dozen complete pterosaurs that should be in phylogenetic analysis… while ignoring the lepidosaurs that gave rise to the ancestors of pterosaurs. Unfortunately, that’s the world academics live in today. They keep trying to not upset the lectures and textbooks from which they make their living. Apparently if academics focus on the details they won’t have to worry about the big picture. No one will ever know the difference if no one points out the elephant in the room.

Averianov AO 2020. Taxonomy of the Lonchodectidae (Pterosauria, Pterodactyloidea). Proceedings of the Zoological Institute RAS. 324 (1): 41–55. doi:10.31610/trudyzin/2020.324.1.41
Bowerbank JS 1846. On a new species of pterodactyl found in the Upper Chalk of Kent (Pterodactylus giganteus). Quarterly Journal of the Geological Society of London. 2: 7–9.
Bowerbank JS 1848. Microscopical observations on the structure of the bones of Pterodactylus giganteus and other fossil animals”. Quarterly Journal of the Geological Society. 4: 2–10.
Martill DM, Smith RE, Longrich N and Brown J 2020. Evidence for tactile feeding in pterosaurs: a sensitive tip to the beak of Lonchodraco giganteus (Pterosauria, Lonchodectidae) from the Upper Cretaceous of southern England. Cretaceous Research
Available online 3 September 2020, 104637 Cretaceous Research
Rodrigues T and Kellner A 2013. Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England. ZooKeys. 308: 1–112. doi:10.3897/zookeys.308.5559


“Kinematics of wings from Caudipteryx to modern birds”: Talori et al. 2018

A new paper without peer-review by Talori, Zhao and O’Connor 2018
seeks to “better quantify the parameters that drove the evolution of flight from non-volant winged dinosaurs to modern birds.”

they employ Caudipteryx, an oviraptorosaur. They correctly state,
Currently it is nearly universally accepted that Aves belongs to the derived clade of theropod dinosaurs, the Maniraptora.” They incorrectly state, “The oviraptorosaur Caudipteryx is a member of this clade and the basal-most  maniraptoran with pennaceous feathers.” In the large reptile tree (LRT, 1269 taxa) oviraptorosaurs nest with therizinosaurus, and more distantly ornithomimosaurs. This clade is separated from bird ancestor troodontids by the Ornitholestes/Microraptor clade.

Figure 1. More taxa, updated tree, new clade names.

Figure 1. Caudipterys is in the peach-colored clade, far from the lineage of birds.

The Talori team
mathematically modeled Caudipteryx with three hypothetical wing sizes, but failed to provide evidence that the Caudipteryx wing was capable of flapping. In all flapping tetrapods the elongation of the coracoid  (or in bats of the clavicle) signals the onset of flapping… and Caudipteryx does not have an elongate coracoid. Rather, it remains a disc.

So, no matter the math, or the accuracy of the mechanical model,
the phylogeny is not valid and the assumption of flapping is inappropriate. It would have been better if they had chosen a troodontid and several Solnhofen birds to test.

Tossing those issues aside,
the Talori team did an excellent job of setting their mechanical model (which could be a troodontid) in a wind tunnel, extracting data from three different wing shapes and presenting their findings. Feathers would have been more flexible than their mold manufactured wings, but the effort is laudable.

Zhao J-S, Talori YS, O’Connor J-M 2018. Kinematics of wings from Caudipteryx to modern birds. [not peer-reviewed] bioRXiv