Earlier one of the the strangest of all bottom-dwelling vertebrates,
Squaloraja (Fig. 1), nested in the LRT with paddlefish (= Polyodon) and goblin sharks (= Mitsukurina), rather than the traditional ratfish (= Chimaera).
a traditional relative of Mitsukurina, Scapanorhynchus (Davis 1887, Woodward 1889, 1899; Figs. 2, 3) nests closer to Squaloraja (Fig. 1) in the paddlefish clade.
(originally Rhinognathus lewisii David 1887; Woodward 1889; NHMUK PV P 4774; Early Cretaceous; 65cm to 1m in length) is widely considered a relative of Mitsukurina, the goblin shark, but here nests closer to Squaloraja. Here the gill basket is much longer, the eyes are midway in size and two dorsal fins are retained.
Squaloraja polyspondyla (Agassiz 1843, Woodward 1866, Early Jurassic) is traditionally considered a relative of Chimaera, but here nests with Scapanorhynchus from the Cretaceous.
This appears to be
another novel hypothesis of interrelationships. If not, please supply the citation so I can promote it.
Agassiz L 1843. Recherches sur les Poissons Fossiles, III (IV), Imprimerie de Petitpierre, Neuchatel, pp. 157-390.
Davis JW 1887. The fossil fishes of the chalk of Mount Lebanon, in Syria. Scientific Transactions of the Royal Dublin Society, 2 (3): 457–636, pl. 14–38.
Woodward AS 1886. On the anatomy and systematic position of the Liassic selachian Squaloraja polyspondyla Agassiz. Proceedings of the Zoological Society of London, 1886: 527–538.
Woodward AS 1889. Catalogue of the Fossil Fishes in the British Museum. Part 1. London: British Museum of Natural History, 1-474.
Woodward AS 1899. Note on Scapanorhynchus, a Cretaceous shark apparently surviving in Japanese seas. Annals and Magazine of Natural History, (Series 7), 3 (18): 487–489.