Caupedactylus ybaka (Kellner 2012) enters the LPT

Kellner (2012, 2013) described
the skull of an Early Cretaceous sinopterid pterosaur, Caupedactylus ybaka (MN 4726-V, Fig. 1). The skull is about forty-six centimetres long. (Hope I got this right this time).

Earlier the same skull was posted online.

Figure x. Caupedactylus in situ and restored by sculptors.

Figure x. Caupedactylus in situ and restored by sculptors. Or a different specimen.

New Tapejarid-Tupuxuarid skull.

Figure 1. New Tapejarid-Tupuxuarid skull now named Caupedactylus.

Bones colorized in this tapejarid / tupuxuarid.

Figure 2. Bones colorized in this tapejarid / tupuxuarid, named Caupedactylus.

Abstract
“A new unusual tapejarid pterosaur from the Early Cretaceous Romualdo Formation (Araripe Basin, Brazil) is described, based on a skull, lower jaw and some postcranial elements. Caupedactylus ybaka gen. et sp. shows the typical high nasoantorbital fenestra of the Thalassodrominae but lacks a palatal ridge, and shares with the Tapejarinae several features, including a downturned rostral end, allowing its allocation to that clade.”

The new skull compared to other tapejarids. Click to enlarge.

Figure 2. Click to enlarge. The rising size of the tapejaridae.

Abstract continues
“Furthermore, the new species differs in having an anteriorly and posteriorly expanded premaxillary sagittal crest, the lacrimal process of the jugal strongly inclined, and a slit-like postpalatine fenestra, among other characters. The region of the left jugal-quadratojugal-quadrate shows a pathology that is likely the result of an infection. The lateral surface of the premaxillary crest presents grooves that were interpreted in other pterosaurs as impressions of blood vessels, corroborating growing evidence that cranial crests could have been involved in thermoregulation.”

“Also, the new species has a well-preserved palate with a large palatine forming the anterior region of the choanae and the postpalatine fenestra and a secondary subtemporal fenestra. Since the latter has been regarded as unique to non-pterodactyloids, its occurrence in Caupedactylus demonstrates that the evolution of palatal region in pterosaurs is more complex than previously thought.”

Perhaps to no one’s surprise, this specimen nested in 2013 in the large pterosaur tree (LPT) between Sinopterus dongi and Tupandactylus.


References
Campos HBN and Headden JA 2013. A review of Tupuxuara deliradamus (Pterosauria, Azhdarchoidea, Thalassodromidae) from the Early Cretaceous Romualdo Formation of Brazil. International Symposium on Pterosaurs – Rio Ptero 2013.
Elgin RA 2015. Paleobiology, Morphology and Flight Characteristics of Pterodactyloid Pterosaurs. Dissertation, University of Heidelberg.
Kellner AWA 2012. A new unusual tapejarid (Pterosauria, Pterodactyloidea) from the Early Cretaceous Romualdo Formation, Araripe Basin, Brazil. Cambridge University Press 103 (3-4) The Full Profession: A Celebration of the Life and Career of Wann Langston, Jr., Quintessential Vertebrate Palaeontologist September 2012 , pp. 409-421.
Kellner AWA 2013. A new unusual tapejarid (Pterosauria, Pterodactyloidea) from the Early Cretaceous Romualdo Formation, Araripe Basin, Brazil. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 103(3-4): 409-421.
Manzig PC et al. (10 co-authors) 2014. Discovery of a Rare Pterosaur Bone Bed in a Cretaceous Desert with Insights on Ontogeny and Behavior of Flying Reptiles. PLoS ONE 9(8): e100005.
Martill DM and Naish D 2006. Cranial Crests Development in the Azhdarchoid Pterosaur Tupuxuara, With Review of the Genus and Tapejarid Monophyly. Palaeontology 49(4): 925-941.

wiki/Caupedactylus
pterosaurheresies.wordpress.com/2013/06/06/tapejarid-or-tupuxuarid/

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