Overlooked jugal on a king mackerel

Short one today. 
As I confess to overlooking the jugal on the skull of the king mackerel, Scomberomorus (Figs. 1, 2)… yet another moment of discovery and correction, putting this puzzle together.

Figure 2. The skull of the king mackerel, Scombermorus cavalla.

Figure 2. The skull of the king mackerel, Scombermorus cavalla. I overlooked the jugal at this point and considered it part of the pterygoid. 

Figure 2. Scomberomorus again, this time with the jugal highlighted in cyan.

Figure 2. Scomberomorus again, this time with the jugal highlighted in cyan.

Reviewing the cladogram
brought this issue to my attention. Usually the data collector informs the data matrix. Here  the cladogram informed the data collector.

Little jugals pasted over the hyomandibular
were also identified on several related taxa.

Scomberomorus cavalla (Cuvier 1829; 60cm) is the extant king mackeral or kingfish. Traditionally assocated with Scomber the mackerel, here it nests with Aphanopus. These taxa are basal to flying fish + swordfish and more distantly, sticklebacks + sea horses. King mackerels are derived from the barracuda, Sphyraena + the mahi-mahi, Coryphaena.


References
Cuvier GCLD 1829. Le Règne Animal distribué d’apres son organisation, pour servir de base a l’histoire naturelle des animaux et d’introduction a l’anatomie comparée. Avec figures dessinées d’après nature. Nouvelle édition, revue et augmentée. Tome V. Suite et fin des Insectes. Par M. Latreille. Déterville & Crochard, Paris, i-xxiv + 556pp.

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