This one caught me by surprise, too.
It came about because I re-examined the details.
the speedy flying fish (Exocoetus, Fig. 1) seems to have little in common with the lethargic ocean sunfish (Mola, Fig. 2) and opah (Lampris, Fig. 3). Traditionally flying fish nest with speedy needlefish, like Tylosurus. The large reptile tree (LRT, 1647+ taxa) adds 25 more steps to move Exocoetus next to Tylosurus.
provide a different hypothesis of interrelationships. You can see a short deep mandible and a deep coracoid in these three taxa (Figs. 1-3), along with a long list of other homologous traits.
All three taxa are derived from
the speedy high-fin amberjack, Seriola rivoliana.
Exocoetus volitans (Linneaus 1758; up to 30cm ) is the extant blue flyingfish, here related to the much larger and nonvolant opah, Lampris (above). Exocoetus travels in schools or schoals. Sometimes they exit the water to avoid predators. Juveniles have a relatively shorter torso. Hatchlings are slow-moving and tiny. Distinctly flying fish and their relatives have a jaw joint directly below the orbit. The coracoid is larger than the scapula, raising and powering the pectoral fins.
Linnaeus C 1758. Systema naturæ per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata.