Mongoose trifles

Herpestes, the Egyptian mongoose, 
 (Linneaus 1758; extant; 48-60cm in length) has large carnassials. Herpestes is a lower, shorter-legged ancestor to the raccoon, Procyon, with a relatively shorter rostrum and longer, lower body. Surprisingly, the postfrontal and postorbital are elongated here.

Figure 1. The Egyptian mongoose, Herpestes, develops a postorbital bar arising from the layered postfrontal and postorbital reappearing in this clade.

Figure 1. The Egyptian mongoose, Herpestes, develops a postorbital bar arising from the layered postfrontal and postorbital reappearing in this clade. The lacrimal and prefrontal are separated here.

Eupleres from Madagascar 
(Doyère 1835) is the extant Western falnouc, a cat-like mongoose from Madagascar. Note the elongate premaxilla, the gracile mandible, the reduced canine and other rodent-like traits. No postfrontal or postorbital appears here.

Figure 2. Eupleres is a Madagascar mongoose with a long, tree-shrew-like skull with a longer premaxilla.

Figure 2. Eupleres is a Madagascar mongoose with a long, tree-shrew-like skull with a longer premaxilla and smaller, more widely-space, primitive teeth. No postfrontal or postorbital appears here.

Despite the differences in these two taxa,
the large reptile tree nests them in the same clade along with Prohesperocyon, the Late Eocene pre-mole, and Talpa the living mole (a member of Carnivora, not Insectivora).

References
Doyére LMF 1835. Notice sur un mammifére de Madagascar, formant le type d’un nouveau genre de la famille des Carnassiers insectivores de M. Cuvier. Ann. Sci. Nat. Zool. 4: 270–283.
Linnaeus C von 1758. Systema naturæ per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata.

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