Wilberg, Turner and Brochu 2019
bring us the “evolutionary structure and timing of major habitat shifts in Crocodylomorpha.”
Wilberg, Turner and Brochu 2019 omitted a long list of basal bipedal crocodylomorpha (Figs. 1, 2), and considered an ingroup, Gracilisuchus, an outgroup. They also considered the unrelated giant rauisuchid, Postosuchus, an outgroup taxon.
Missing from the Wilberg, Turner and Brochu study
are a number of basal Crocodylomorph taxa and their outgroups, as determined by a wide gamut analysis of reptiles (= the large reptile tree = LRT; subset Fig. 1).
Ingroup Crocodylomorpha in the LRT
omitted by Wilberg, Turner and Brochu include: YPM VP 057 103, Pseudhesperosuchus, Carnufex, Trialestes, Lewisuchus, MCZ 4116, Saltopus, Scleromochlus, SMNS 12591, Terrestrisuchus, Tarjadia, Parringtonia, Litargosuchus, Erpetosuchus, SMNS 12352, Pedeticosaurus, Yonghesuchus, Dromicosuchus, Saltoposuchus and Dyoplax.
Outgroups to the clade Crocodylomorpha in the LRT omitted by Wilberg, Turner and Brochu include: Herrerasaurus + Staurikosaurus (at the base of the Dinosauria), PVL 4597 (at the base of the Archosauria = Crocodylomorpha + Dinosauria), Turfanosuchus (proximal outgroup to Archosauria), Decuriasuchus + Pagosvenator (second proximal outgroup to Archosauria).
Simply adding taxa
solves all these problems and documents a more gradual accumulation of derived traits and major habitat shifts.
Wilberg EW, Turner AH and Brochu CA 2019. Evolutionary structure and timing of major habitat shifts in Crocodylomorpha. Nature.com/scientificreports DOI:10.1038/s41598-018-36795-1