More data on the Aetosauroides skull

Seven years ago Aetosauroides nested between
Ticinosuchus and Aetosaurus at the base of the Aetosauria in the large reptile tree (LRT, 1269 taxa) based on a line drawing of the skull lacking several parts (Fig. 1). We first talked about this nesting here in 2011.

Today
Brust et al. 2018 bring more complete data on the skull of Aetosauroides (Fig. 1). As far as the nesting goes, the previous line drawing was good enough, despite the missing parts and cartoonish tracing.

Figure 1. Aetosauroides skull UFSM 11505 from Brust et al. 2018 with color applied to bones here. Small inset shows best previous data.

Figure 1. Aetosauroides skull UFSM 11505 from Brust et al. 2018 with color applied to bones here. Small inset shows best previous data.

Unfortunately,
Brust et al. excluded Ticinosuchus (Figs. 2, 3) from the cladogram. Instead they employed the unrelated Revueltosaurus and Postosuchus as outgroup taxa. As readers know by now, taxon exclusion has hampered many studies in the past and, apparently, this continues into the present.

Figure 2. Aetosaur skulls compared to Ticinosuchus, the long-sought outgroup to this clade.

Figure 2. Aetosaur skulls compared to Ticinosuchus, the long-sought outgroup to this clade.

Vjushkovia, Ticinosuchus and the base of the Stagonolepidae (aetosaurs)

Figure 3. Vjushkovia, Ticinosuchus and the base of the Stagonolepidae (aetosaurs)

The new data for Aetosauroides
was added to the LRT and the tree topology did not change. Revueltosaurus continues to nest with Fugusuchus, far from aetosaurs.

Figure 4. Subset of the LRT focusing on the Euarchosauriformes, including the Aetosauria.

Figure 4. Subset of the LRT focusing on the Euarchosauriformes, including the Aetosauria.

References
Brust ACB, Desojo JB, Schultz CL, Paes-Neto VD and Da-Rosa AAS 2018. Osteology of the first skull of Aetosauroides scagliai Casamiquela 1960 (Archosauria: Aetosauria) from the Upper Triassic of southern Brazil (Hyperodapedon Assemblage Zone) and its phylogenetic importance. https://doi.org/10.1371/journal.pone.0201450

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