The skull of the smooth chameleon,
Chamaeleo laevigatus (Figs. 1, 2), has two extra holes in the anterodorsal plane of its rostrum (Fig. 1). Despite appearances, the holes visible in top view are not nostrils.
The Chamaeleo rostrum
is angled at about 50º from the jawline. Given just the skull, you might think those openings in dorsal view are nostrils. With skin and scales on (Fig. 2), the nostrils are located on the lateral plane, as in other chameleons, like Trioceros (Fig. 1), surrounded by traditional circumnarial bones.
Diaz and Trainer 2015 published
some nice images of chameleon hands and feet, colorized here (Fig. 3) for additional clarity. The metacarpals and metatarsals are the bones that radiate. The phalanges are all vertical here.
Diaz RE Jr. and Trainor PA 2015. Hand/foot splitting and the ‘re-evolution’ of mesopodial skeletal elements during the evolution and radiation of chameleons. BMC Evolutionary Biology201513:184.