Dr. Michael Coates for sending a pdf of his 1995 study of Tulerpeton. From the improved data I was able to make new reconstructions of the manus and pes. The differences shift the nesting of Tulerpeton to the last common ancestor of all reptiles (= amniotes). replacing Gephyrostegus bohemicus, the taxon that held that node in the large reptile tree (LRT) for the last six years.
In the new reconstruction
only the manus retained 6 digits, with the lateral sixth digit a vestige. The pes has a new reconstruction with only 5 digits, very much in the pattern of Gephyrostegus bohemicus. Both have five phalanges on digit 5. In the new reconstructions all of the PILs (Peters 2000) line up in sets.
Lebedev and Coates report:
“A cladistic analysis indicates that Tulerpeton is a reptilomoprh stem-group amniote and the earliest known crown-group tetrapod. The divergence of reptilomorphs from batrachomorphs (frogs and kin) occurred before the Devonian Carboniferous boundary. Polydactyly persisted after the evolutionary divergence of the principal lineages of living tetrapods. Tulerpeton was primarily air-breathing.” They did not test Silvanerpeton, Gephyrostegus, Eldeceeon or Urumqia, which all now nest as proximal kin to Tulerpeton.
Manual digit 6 is present as a vestige. An anocheithrum (small bone atop the cleithrum) is present. Metatarsal 1 in Tulerpeton is the largest in the set. The posterior ilium rises. The femur has a large, sharp, fourth (posterior) trochanter.
on Tulerpeton are also found similar in size and number are also found in related taxa.
and digital graphic segregation AND reconstruction AND comparative anatomy all contributed to the new data scores. As usual, I have not seen the specimen, but I did add it to a large gamut data matrix, the likes of which are not typically employed.
Coates MI and Ruta M 2001 (2002). Fins to limbs: What the fossils say. Evolution & Development 4(5): 390–401.
Lebedev OA 1984. The first find of a Devonian tetrapod in USSR. Doklady Akad. Navk. SSSR. 278: 1407–1413.
Lebedev OA and Clack JA 1993. Upper Devonian tetrapods from Andreyeva, Tula Region, Russia. Paleontology36: 721-734.
Lebedev OA and Coates MI 1995. postcranial skeleton of the Devonian tetrapod Tulerpeton curtum Lebedev. Zoological Journal of the Linnean Society. 114 (3): 307–348.
Peters D 2000. Description and Interpretation of Interphalangeal Lines in Tetrapods. Ichnos, 7: 11-41