A few years ago
a rather complete lizard skull (BGS GSb581) was described (Evans et al. 2012) from the Purbeck Limestone (Late Jurassic to Early Cretaceous) of England. It was originally excavated more than a century ago and assigned to the genus, Paramacellodus. Evans et al. renamed it Purbicella. Their cladistic analysis nested Purbicella with Lacertoidea: (Lacertidae (including Acanthodactylus), Teiidae (including Tupinambus), Gymnophthalmidae (including Gymnophthalmus), and the burrowing Amphisbaenia (including Amphisbaena)), not Paramacellodus, which nested with skinks. Evans et al. based their nesting on a partial data matrix of Conrad (2008).
The large reptile tree nested Purbicella between Acanthodactylus and Liushusaurus. The large reptile tree recovered the above listed ‘lacertoid’ taxa as members of a paraphyletic clade, some preceding Purbicella in various clades and others succeeding it.
While Purbicella is Late Jurassic/Early Cretaceous, it must have had its origins much earlier, in the Late Carboniferous, because a descendant taxon, Ascendonanus, is Early Permian.
Conrad JL 2008. Phylogeny and systematics of Squamata (Reptilia) based on morphology. Bulletin of the American Museum of Natural History 310:1–182.
Evans SE, Jones MEH and Matsumto R 2012. A new lizard skull from the Purbeck Limestone Group (Lower Cretaceous) of England. Bull. Soc. géol. France, 2012, t. 183(6):517-524.