More wing and a foot added to Scaphognathus

The holotype of Scaphognathus (Goldfuss 1830) GPIB 1304, Fig. 1) is gorgeous, but incomplete. The posterior pelvis and tail are missing. So is the distal wing phalanx.

Figure 1. The holotype of Scaphognathus.

Figure 1. The holotype of Scaphognathus.

Earlier I thought the distal tibia and foot were missing, but a closer look at what remains of the left tibia left me thinking the pedal elements were all present, mixed up with the gastralia. I wouldn’t have confidence saying this except the reconstruction of the pedal elements is very much like those of other Scaphognathus specimens, like n110 and the Maxberg specimen.

Figure 2. Scaphognathus pedal elements highlighted in color. These are difficult to distinguish from gastralia.

Figure 2. Scaphognathus pedal elements highlighted in color. These are difficult to distinguish from gastralia. The reconstruction resembles the pes of other Scaphognathus specimens.

So we may have something here.

As a side note, one more wing phalanx is also added to the wing. Most of the left wing is hidden by the antebrachium and pelvis. I missed that earlier. So, here are the images.

Figure 3. Three proximal wing phalanges for Scaphognathus in red, green and blue.

Figure 3. Three proximal wing phalanges for Scaphognathus in red, green and blue.

And a new reconstruction puts it all together.

Figure 3. New reconstruction of Scaphognathus with the new foot and wing phalanges added.

Figure 3. New reconstruction of Scaphognathus (n109) with the new foot and wing phalanges added. If I had imagined a foot, I would have made it larger. The metatarsus was found separated, but sister taxa have a compressed metatarsus. It is also possible that more of the proximal metatarsus is missing off the edge of the matrix, but I don’t think so. More of pedal 1.1 may be present beneath the femur to match sister taxa.

Hopefully a new basal Scaphognathus specimen will come along with a foot and wing in clear view. It is entirely possible that this “foot” is created from broken and half-sunken gastralia, but at this point, it’s worth presenting.

References
Goldfuss GA 1830. Pterodactylus crassirostris. Isis von Oken, Jena pp. 552–553.
Wellnhofer P 1975a. Teil I. Die Rhamphorhynchoidea (Pterosauria) der Oberjura-Plattenkalke Süddeutschlands. Allgemeine Skelettmorphologie. Paleontographica A 148: 1-33.1975b. Teil II. Systematische Beschreibung. Paleontographica A 148: 132-186. 1975c. Teil III. Paläokolgie und Stammesgeschichte. Palaeontographica 149: 1-30.

wiki/Scaphognathus

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